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The O104:H4 isolates, including the outbreak isolates, clustered into a single, closely related clade — again, with the exception of the C35-10 isolate. The four isolates from the German outbreak (TY2482, C227-11, LB226692, and H112180280), three of which were sequenced by other groups, are particularly closely related. The Lambdalike Phage Containing the Stx2 Genes A polymerase-chain-reaction assay revealed that the outbreak isolates contained genes encoding Shiga toxin 2, a toxin that inhibits protein synthesis and causes the hemolytic–uremic syndrome.
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The pAA C227-11 plasmid encodes a number of enteroaggregative E. Coli–specific virulence factors (, and Tables 8 and 9 in the ), such as aggR and genes that it regulates, including AggR, AatPA–D, Aap, AggA–D, and SepA, and thus presumably is critical for C227-11 pathogenicity. Discussion The outbreak clone is a dramatic example of gene acquisition by means of lateral transfer that resulted in an accretion of synergistic virulence factors. By all molecular definitions, it is an enteroaggregative E. Coli strain but one that has acquired a Shiga-toxin–encoding phage — a hallmark of Shiga-toxin–producing E.
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The O104:H4 isolates, including the outbreak isolates, clustered into a single, closely related clade — again, with the exception of the C35-10 isolate. The four isolates from the German outbreak (TY2482, C227-11, LB226692, and H112180280), three of which were sequenced by other groups, are particularly closely related. The Lambdalike Phage Containing the Stx2 Genes A polymerase-chain-reaction assay revealed that the outbreak isolates contained genes encoding Shiga toxin 2, a toxin that inhibits protein synthesis and causes the hemolytic–uremic syndrome.


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